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Because the active (non-protonated) Meta II conformation absorbs at 378 nm, it appears that most of the chromophore in the triple mutant is bound via a non-protonated Schiff-based linkage

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A., Ito, T., Michlek, M., Sugimoto, S., Kita, H., Chijimatsu, R., Ukon, Y., Kushioka, J., Okada, R., Kamatani, T., Hashimoto, K., Tateiwa, D., Tsukazaki, H., Nakagawa, S., Takenaka, S., Makino, T., Sakai, Y., Necas, D., Kaito, T