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Ethics declarations Ethical approval All animal studies followed the ethical policies and guidelines recommended by ARRIVE (Animal Research: Reporting of In Vivo Experiments) and National Research Councils Guide for the Care and Use of Laboratory Animals and were approved by the Ethics Committee of the Animal Experimentation of Jinan University (Approval No.: 20230901-0012)

doi: 10.1038/s41598-018-34803-y 47 ScalcoEFiorinoCCattaneoGMSanguinetiGRizzoG

ADP, average daily pain

Plating and treatment After two passages, 30.000 cells were plated in 24-well plates with coverslips pre-coated with poly-L-ornithine (sigma, P4957)