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Interestingly, the categories enriched in the upregulated clusters are related to phagosome maturation, RNA and translation, but importantly multiple metabolism pathways including citric acid cycle, cholesterol metabolism, pentose phosphate pathway, electron transport chain, and glutathione metabolism

doi:10.1002/14651858.CD011003.pub2

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The copper-bound complex delivers Cu to copper-dependent enzymes such as lysyl oxidase, which catalyzes collagen and elastin cross-linking