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3.2 Increasing AsA content by modulating its biosynthesis is not easy Over the years, virtually all the genes involved in the Smirnoff-Wheeler pathway have been overexpressed under the control of constitutive promoters to obtain higher AsA content in different plant species, but in almost all cases the actual results were far below the expected (Strobbe et al., 2018), the only valuable exception being GGP ( Table 2 )