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Wilson, J.R

The first line of defence is the plant cell wall and cuticular barrier, which not only provides a protective shield against physical damage by animals, insects, abiotic stresses (e.g., freeze-thawing) and bacterial and fungal pathogens that is not simply passive but transmits danger signals within the plant as immunogenic factors

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In addition to these enzymatic antioxidants, sperm and seminal plasma contain several non-enzymatic antioxidants that provide further protection against oxidative stress